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An. * Address correspondence to george.haughn@ubc.ca. The author responsible for distribution of materials integral to the findings presented in this article in accordance with the policy described in the Instructions for Authors (www.plantphysiol.org) is: George W. Haughn (george.haughn@ubc.ca). A.Y.-L.T. designed the research; A.Y.-L.T., T.K., and J.R. performed the research; A.Y.-L.T., L.J.F.,

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