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E of patterning many developing tissues. More recently, Hh signaling has been implicated in both Dmel and mouse adult stem cell maintenance and human neoplastic disease (Bale, 2002; Ihrie et al., 2011; Jiang and Hui, 2008; Rojas-R s et al., 2012; Scales and de Sauvage, 2009; Varjosalo and Taipale, 2008). While a number of papers have investigated Hh extracellular movement and downstream signaling,

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