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Progression) we focus on the apical/basal transport of Hh in developing photoreceptors in healthy and disease models, as this is particularly difficult to observe and quantify in other established models of Hh in development (Jiang and Hui, 2008; Yazawa et al., 2009). More broadly, this method could be adapted to additional neurodegenerative disease models (e.g. in the Dmel eye) and should enable

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